S.C. Weeks Bibliography


Click on Outlined Names for an abstract or pdf of each paper.

Calabrese, A, C McCullough, B Knott and SC Weeks. Environmental characteristics of ephemeral rock pools explain local abundances of the clam shrimp, Paralimnadia badia (Branchiopoda: Spinicaudata: Limnadiidae). Journal of the Royal Society of Western Australia (in press).


Weeks, SC, JS Brantner, TI Astrop, DW Ott and N Rabet. 2013. The evolution of hermaphroditism from dioecy in crustaceans: Selfing hermaphroditism described in a fourth Spinicaudatan genus. Evolutionary Biology DOI 10.1007/s11692-013-9265-0.


Brantner, J. S. , D. W. Ott, R. J. Duff, L-O Sanoamuang, GP Simhachalam, KK Subhash Babu, and SC Weeks. 2013. Androdioecy and Hermaphroditism in Five Species of Clam shrimp (Crustacea: Branchiopoda: Spinicaudata) From India and Thailand. Invert. Biol. 132:27-37.


Brantner, J. S. , D. W. Ott, R. J. Duff, J.I. Orridge, J. R. Waldman and S.C. Weeks. 2013. Evidence of selfing hermaphroditism in the clam shrimp Cyzicus gynecia (Branchiopoda, Spinicaudata). J. Crust. Biol. 33:184-190.

Weeks, S.C. 2012. The role of androdioecy and gynodioecy in mediating evolutionary transitions between dioecy and hermaphroditism in the Animalia. Evolution 66:3670-3686.

Astrop, TI , LE Park, B Brown and SC Weeks. 2012. Sexual discrimination at work: Spinicaudatan ‘Clam Shrimp’ (Crustacea: Branchiopoda) as a model organism for the study of sexual system evolution. Palaeontologia Electronica 15.2.20A.

Rogers, DC, N Rabet, and SC Weeks. 2012. A revision of the extant genera of the Limnadiidae (Branchiopoda, Spinicaudata) J. Crust. Biol. 32:827-842.

Benvenuto, C, and SC Weeks. 2012. Intersexual conflict during mate guarding in an androdioecious crustacean. Behav. Ecol. 23:218–224.

Benvenuto C,and SC Weeks. 2011. Mate guarding behavior in clam shrimp: The influence of mating system on intersexual conflict. Behav. Ecol. Sociobiol. 65:1899-1907.

Weeks SC, TF Sanderson , BF Wallace, and B Bagatto. 2011. Behavioral cost of reproduction in a freshwater crustacean (Eulimnadia texana). Ethology 117:880-886.

Weeks SC, C Benvenuto , TF Sanderson, and RJ Duff. 2010. Sex Chromosome Evolution in the Clam Shrimp, Eulimnadia texana. J. Evol. Biol. 23:1100-1106.

Rogers DC, SC Weeks, and WR Hoeh. 2010. A new species of Eulimnadia (Crustacea; Branchiopoda; Diplostraca; Spinicaudata) from North America. Zootaxa 2413:61-68.

Weeks SC, EG Chapman , DC Rogers, DM Senyo, and WR Hoeh. 2009. Evolutionary transitions among dioecy, androdioecy and hermaphroditism in limnadiid clam shrimp (Branchiopoda: Spinicaudata). J. Evol. Biol. 22:1781-1799.

Benvenuto C, B Knott and SC Weeks. 2009. Mate guarding behavior in clam shrimp: A field approach. Behav. Ecol. 20:1125-1132.

Weeks SC, SK Reed , DW Ott, and F. Scanabissi. 2009. Inbreeding effects on sperm production in clam shrimp (Eulimnadia texana). Evol. Ecol. Res. 11:125-134.

Weeks SC. 2009. Can males successfully invade hermaphroditic populations of clam shrimp (Eulimnadia texana)? Current Science 96:98-102.

Weeks SC, TF Sanderson , M Zofkova, and B Knott. 2008. Breeding systems in the clam shrimp family Limnadiidae. Invertebrate Biology 127:336-349.

Brendonck L, DC Rogers, J Olesen, S Weeks, R Hoeh. 2008. Global diversity of large branchiopods (Crustacean; Branchiopoda) in freshwater. Hydrobiologia 595:167-176.

Weeks SC, and C Benvenuto. 2008. Mate guarding in the androdioecious clam shrimp Eulimnadia texana: male assessment of hermaphrodite receptivity. Ethology 114:64-74.

Duff RJ, C Benvenuto, T Branch, and SC Weeks. 2007. DNA extraction from cysts of the clam shrimp Eulimnadia texana (Crustacea, Branchiopoda, Spinicaudata). Journal of Crustacean Biology 27:154-157.

Weeks SC, M Zofkova, and B. Knott. 2006. Limnadiid Clam Shrimp Distribution in Australia (Crustacea: Branchiopoda: Spinicaudata). Journal of the Royal Society of Western Australia 89:155-161.

Weeks SC, SK Reed, M Cesari, and F Scanabissi. 2006. Production of intersexes and the evolution of androdioecy in the clam shrimp Eulimnadia texana (Crustacea, Branchiopoda, Spinicaudata). Invertebrate Reproductive Development 49:113-119.

Weeks SC, C Benvenuto, and SK Reed. 2006. When males and hermaphrodites coexist: A review of androdioecy in animals. Integrative and Comparative Biology 46:449-464.

Hoeh WR , ND Smallwood , DM Senyo, EG Chapman, and SC Weeks. 2006. Evaluating the monophyly of Eulimnadia and the Limnadiinae (Branchiopoda: Spinicaudata) using DNA sequences. Journal of Crustacean Biology 26:182-192.

Weeks SC, TF Sanderson, SK Reed, M Zofkova, B Knott, U Balaraman, G Pereira, DM Senyo, WR Hoeh. 2006. Ancient Androdioecy in the Freshwater Crustacean Eulimnadia . Proceedings of the Royal Society of London, Series B 273:725-734 .

Scanabissi F, M Cesari, SK Reed, SC Weeks. 2006. Ultrastructure of the male gonad and male gametogenesis in Eulimnadia texana (Packard, 1871)(Crustacea, Branchiopoda, Spinicaudata). Invertebrate Biology 125:117-124.

Weeks, SC, RT Posgai, M Cesari, and F Scanabissi. 2005. Androdioecy inferred in the clam shrimp Eulimnadia agassizii (Spinicaudata: Limnadiidae). Journal of Crustacean Biology 25:323-328.

Duff, RJ WR Hoeh, DG Cook and SC Weeks. 2004. Isolation and characterization of thirteen polymorphic microsatellite loci from the clam shrimp Eulimnadia texana (Crustacea: Spinicaudata). Molecular Ecology Notes 4:397-399.

Weeks, SC. 2004. Seven generations of inbreeding does not purge inbreeding depression in the clam shrimp, Eulimnadia texana. Journal of Evolutionary Biology 17:475-484.


Weeks, SC and RL Bernhardt. 2004. Maintenance of androdioecy in the freshwater shrimp, Eulimnadia texana: Field estimates of inbreeding depression and relative male survival. Evolutionary Ecology Research 6:227-242.

Weeks SC, CL Marquette, and E Latsch. 2004. Barriers to outcrossing success in the primarily self-fertilizing clam shrimp, Eulimnadia texana (Crustacea, Branchiopoda). Invertebrate Biology 123:146-155.

Weeks, SC and RJ Duff. 2002. A Genetic Comparison of Different Populations of Clam Shrimp in the genus Eulimnadia. Hydrobiologia 486:295-302.

Weeks, SC, V Marcus, R Salisbury, and D Ott. 2002. Cyst Development in the Conchostracan shrimp, Eulimnadia texana. Hydrobiologia 486:289-294.

Hollenbeck VG, N Zucker, W Gould and SC Weeks. 2002. Maintenance Of Androdioecy In The Freshwater Shrimp Eulimnadia texana: Sexual Encounter Rates And Outcrossing Success. Behav. Ecol. 13:561-570.

Naida Zucker, Gabriela A. Aguilar, Stephen C. Weeks,  and L. Garner McCandless. 2002. Variation in reproductive cycle between selfing and outcrossing hermaphrodites in an androdioecious desert shrimp. Invertebrate Biology 121:66-72.

Weeks, SC, J Hutchison, and N Zucker. 2001. Maintenance of androdioecy in the freshwater shrimp, Eulimnadia texana: Do hermaphrodites need males for complete fertilization? Evolutionary Ecology 15:205-221.

Gray, M and SC Weeks. 2001. A comparison of dietary use patterns of clonal and sexual fish (Poeciliidae: Poeciliopsis). Can. J. Fish. Aquat. Sci. 58:1313-1318.

Zucker, N, B Stafki and SC Weeks. 2001. Maintenance of androdioecy in the freshwater shrimp, Eulimnadia texana: Relative longevity of males to hermaphrodites. Can. J. Zool. 79:393-401.

Weeks, SC, BR Crosser, and MM Gray. 2001. Relative fitness of two hermaphroditic mating types in the androdioecious clam shrimp, Eulimnadia texana. Journal of Evolutionary Biology 14:83-94.

Medland, VL, N Zucker, and SC Weeks. 2000. Maintenance of androdioecy in the freshwater shrimp, Eulimnadia texana Parker: Encounters between males and hermaphrodites are not random. Ethology 106:839-848.

Weeks, SC, BR Crosser, B Bennett, MM Gray, and N Zucker. 2000. Maintenance of androdioecy in the freshwater shrimp, Eulimnadia texana: Variation in inbreeding depression between two populations.  Evolution 54:878-887.

Weeks, SC, BR Crosser, MM Gray, JA Matweyou, and N. Zucker. 2000. Is there Sperm Storage in the Clam Shrimp Eulimnadia texana? Invertebrate Biology 119:215-221.

Weeks, SC, and N Zucker. 1999. Rates of Inbreeding in the Clam Shrimp Eulimnadia texana. Can. J. Zool. 77:1402-1408.

Stockwell, CA and SC Weeks. 1999. Translocations and altered evolutionary trajectories in recently established populations of the western mosquitofish (Gambusia affinis) Animal Conservation 2:103-110.

Weeks, SC, V Marcus, and B Crosser. 1999. Inbreeding Depression in a Self-Compatible, Androdioecious Crustacean, Eulimnadia texana. Evolution 53:472-483.

Belk, D, G Mura and SC Weeks. 1998. Untangling confusion between Eubranchipus vernalis and Eubranchipus neglectus (Branchiopoda, Anostraca). Journal of Crustacean Biology 18:147-152.

Weeks, SC and V Marcus. 1997. A survey of the branchiopod crustaceans of Ohio. Ohio Journal of Science 97:86-89.

Marcus, V and SC Weeks. 1997. The effects of pond duration on the life history traits of an ephemeral pond crustacean, Eulimnadia texana. Hydrobiologia, 359:213-221.

Weeks, SC and GK Meffe. 1997. "Grandfather Effects" on offspring size in the Eastern Mosquitofish, Gambusia holbrooki. Copeia 1997:869-874.

Weeks, SC, V Marcus and S Alvarez. 1997. Notes on the life history of the clam shrimp Eulimnadia texana. Hydrobiologia 359:191-197.

Weeks SC. 1996. A reevaluation of Red Queen model for the maintenance of sex in a clonal-sexual fish complex (Poeciliidae: Poeciliopsis). Canadian Journal of Fisheries and Aquatic Sciences 53:1157-1164.

Weeks SC. 1996. The hidden cost of reproduction: reduced energetic intake caused by spatial constraints in the body cavity. Oikos 75:345-349.

Weeks SC and GK Meffe. 1996. Quantitative genetic and optimality analyses of life-history plasticity in the eastern mosquitofish, Gambusia holbrooki. Evolution 50:1358-1365.

Meffe, G.K., S.C. Weeks, P. Mulvey, and K.L. Kandl. 1995. Genetic differences in thermal tolerance of mosquitofish from ambient and thermally elevated ponds. Canadian Journal of Fisheries and Aquatic Sciences 52:2704-2711.

Weeks, S.C. 1995. Comparisons of life-history traits between clonal and sexual fish (Poeciliopsis: Poeciliidae) raised in monoculture and mixed treatments. Evolutionary Ecology 9:258-274.

Heulett ST, SC Weeks and GK Meffe. 1995. Lipid dynamics and growth relative to resource level in juvenile eastern mosquitofish (Gambusia holbrooki: Poeciliidae). Copeia 1995:97-104.

Weeks SC and OE Gaggiotti. 1993. Patterns of offspring size at birth in clonal and sexual strains of Poeciliopsis (Poeciliidae). Copeia 1993:1003-1009.

Weeks SC. 1993. The effects of recurrent clonal formation on clonal invasion patterns and sexual persistence: A Monte Carlo simulation of the frozen niche-variation model. The American Naturalist 141:409-427.

Sassaman C and SC Weeks. 1993. The genetic mechanism of sex determination in the Conchostracan shrimp Eulimnadia texana. The American Naturalist 141:314-328.

Weeks SC. 1993. Phenotypic plasticity of life-history traits in clonal and sexual fish (Poeciliopsis) at high and low densities. Oecologia 93:307-314.

Connor A, KM Weiss and SC Weeks. 1993. Evolutionary models of quantitative disease risk factors. Human Biology 65:917-940.

Weeks SC, OE Gaggiotti, R Schenck, K Spindler, and RC Vrijenhoek. 1992. Feeding behavior in sexual and clonal strains of Poeciliopsis. Behavioral Ecology and Sociobiology 30:1-6.

Weeks SC and JM Quattro. 1991. Life-history plasticity under resource stress in a clonal fish (Poeciliidae: Poeciliopsis). Journal of Fish Biology 39:485-494.

Quattro JM and SC Weeks. 1991. Correlations of egg size and energetic content in sexual and clonal biotypes of Poeciliopsis. Journal of Fish Biology 38:331-334.

Weeks SC and C Sassaman . 1990. Competition in phenotypically variable and uniform populations of the tadpole shrimp Triops longicaudatus (Notostraca: Triopsidae). Oecologia 82:552-559.

Weeks SC. 1990. Life-history variation under varying degrees of intraspecific competition in the tadpole shrimp Triops longicaudatus (LeConte). Journal of Crustacean Biology 10:498-503.

Wetherington JD, SC Weeks, KE Kotora, and RC Vrijenhoek. 1989. Genotypic and environmental components of variation in growth and reproduction of fish hemiclones (Poeciliopsis: Poeciliidae). Evolution 43:635-645.


Colburn EA, SC Weeks, SK Reed. 2007. Diversity and Ecology of Vernal Pool Invertebrates. pp. 105 – 126 in: Science and Conservation of Vernal Pools in Northeastern North America, Calhoun A and P DeMaynadier, eds. CRC Press.

Wood JW, SC Weeks, GR Bentley and KM Weiss. 1994. Human population biology and the evolution of aging. Pp. 19-75 in: Crews DE and RM Garruto (eds.) Biological Anthropology and Aging. Perspectives on Human Variation Over the Life Span. Oxford University Press, New York, NY.


Weeks, SC, C Benvenuto, SK Reed, RJ Duff, ZH Duan, and JR Pannell. A Field Test of two Models for the Maintenance of Androdioecy in the Freshwater Shrimp, Eulimnadia texana. (in prep).

Weeks, SC, J Brantner, TI Astrop, D Ott, and N Rabet. Evolution of Breeding Systems in the Limnadiidae: Selfing Hermaphroditism described in new genus Calalimnadia. (in prep).


Weeks SC. 2006. Under the flashlight: Clam shrimp. Vernalis 9:1-6.

Last Updated : 2-4-14

E-mail address : SCWeeks@uakron.edu.